Heyer, W. R. (1978). Systematics of the fuscus group of the frog genus leptodatylus (amphibia,leptodactylidae. Natural History Museum of Los Angeles County.
Chicago-referens (17:e uppl.)Heyer, W. Ronald. Systematics of the Fuscus Group of the Frog Genus Leptodatylus (amphibia,leptodactylidae. California: Natural History Museum of Los Angeles County, 1978.
MLA-referens (9:e uppl.)Heyer, W. Ronald. Systematics of the Fuscus Group of the Frog Genus Leptodatylus (amphibia,leptodactylidae. Natural History Museum of Los Angeles County, 1978.
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